Three new species of salamander have been discovered in a remote forest reserve in Costa Rica. They were among some 5,000 plants and animals recorded by scientists from London's Natural History Museum during three expeditions to Central America.
Two species are nocturnal, while the third is a dwarf variety, growing to little longer than a thumbnail.
The three new finds bring the number of Costa Rican salamanders known to science to a total of 43.
Salamanders eat insects and worms, and live in water or in moist areas. They usually feed at night and hide during the day, often hibernating during the winter.
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